ここは角竜に対する理解をより深めるための場所。
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| Bistro Ceratopsia | Ceratopsian Cladogram |
| 角竜の記載カレンダー | |
| 角竜名鑑 | |
| 角竜の見れる博物館 Ceratopsians in Museum |
|
| 手持ち角文献 Bibliography which I have 雑記 → Recent Study |
|
| 角竜の形質 Diagnosis |
管理人が思うクラドグラム
クリックすれば各タクサのキャラクターが出てくるように…頑張ります
―o ・DINOSAURIA Owen, 1842
|― ・SAURISCHIA Seeley, 1887
`―o ・ORNITHISCHIA Seely, 1887
`―o ・Fabrosauridae
`―o ・GENASAURIA
|―o ・THYREOPHORA Nopsca, 1915
`―o ・CERAPODA Sereno
|― ・Heterodontosauridae Romer, 1966
`―+― ・ORNITHOPODA Marsh, 1871
`―o ・MARGINOCEPHALIA Sereno, 1986
|― ・PACHYCEPHALOSAURIA Maryanska & Osmolska, 1982
`― ・CERATOPSIA Marsh, 1890
↑You et al., 2003によれば、HeterodontosaurusはMarginocephaliaと姉妹群となったらしい。
CERATOPSIAの中での分岐分析・・・かなり混乱してます。まずは大まかな分類から・・・
CERATOPSIA Marsh, 1890
Yinlong dowsoni Xu, Forster, Clark & Mo, 2006
Chaoyangsaurus youngi Zhao, Cheng & Xu, 1999
PSITTACOSAURIA Osborn, 1923
Psittacosauridae Osborn, 1923
Psittacosaurus Osborn, 1923
P. sinensis Young, 1958
P. meileyingensis Sereno, Chao, Cheng & Rao, 1988
P. mongoliensis Osborn, 1923
P. xinjiangensis Sereno & Chao, 1988
P. neimongoliensis Russell & Zhao, 1996
P. ordosensis Russell & Zhao, 1996
P. sibiricus Voronkevich & Averianov, 2000
P. lujiatunensis Zhou, Gao, Fox and Chen 2006
Hongshanosaurus houi You, Xu & Wang, 2003
NEOCERATOPSIA Sereno, 1991
Liaoceratops yanzigouensis Xu, Makovichy, Wang, Norell & You, 2002
Yamaceratops dorngobiensis Makovicky & Norell
Archaeoceratops oshimai Dong & Azuma, 1997
Leptoceratopidae
Asiaceratops salsopaludalis Nesov, Kaznyshkina & Cherepanov, 1989
Montanoceratops cerorhynchus Sternberg, 1951
Udanoceratops tschizhovi Kurzanov, 1992
Leptoceratops gracilis Brown, 1914
CORONOSAURIA Sereno, 1986
Protoceratopidae
Bagaceratops rozhdestvenskyi Maryanska & Osmolska, 1975
Breviceratops kozlowskii Maryanska & Osmolska, 1975
Graciliceratops mongoliensis Sereno, 2000
Protoceratops andrewsi Granger & Gregory, 1923
CERATOPSOIDEA Hay, 1902
CERATOPSOMORPHA
Zuniceratops christopheri Wolfe & Kirkland, 1998
Turanoceratops tardabilis Nesov, Kaznyshkina & Cherepanov, 1989
Ceratopsidae Marsh, 1890
Centrosaurinae Lambe, 1915
Albertoceratops nesmoi Ryan, 2007
Avaceratops lammersi Dodson, 1986
Brachyceratops montanensis Gilmore, 1914
Monoclonius crassus Cope, 1876
Centrosaurini
Centrosaurus apertus Lambe, 1904b
Styracosaurus Lambe, 1913
S. albertensis Lambe, 1913
S. ovatus Gilmore, 1930
Pachyrhinosaurini sensu Sampson, 1995
Einiosaurus procurvicornis Sampson, 1995
Achelousaurus horneri Sampson, 1995
Pachyrhinosaurus Sternberg, 1950
P. sp.
P. canadiensis Sternberg, 1950
Chasmosaurinae Lambe, 1915
Chasmosaurini
Pentaceratops sternbergii Osborn, 1923
Agujaceratops mariscalensis Lucas, Sullivan & Hunt, 2006
Chasmosaurus Lambe, 1914
C. russelli Sternberg, 1940
C. belli Lambe, 1914
C. irvinensis Holmes, Forster, Ryan & Shepherd, 2001
Anchiceratops ornatus Lambe, 1914
Arrhinoceratops brachyops Parks, 1925
Torosaurus latus Marsh, 1891
Triceratops horridus Marsh, 1889
1872 Agathaumas sylvestris Cope →AMNH 4000；腸骨と脊椎の一部
1874 Polyonax
mortuarius Cope →断片的角芯
1876 Dysganus bicarinatus Cope →AMNH
397；歯
1876 Dysganus encaustus Cope →AMNH
5739；歯でDysganusの模式種
1876 Dysganus haydenianus Cope →AMNH
573；歯
1876 Dysganus peiganus Cope →AMNH 397；歯
1876 Monoclonius crassus Cope →AMNH 3998；頭蓋等各部分
1887
Bison alticornis Marsh →USNM 1871；上眼窩角芯
1888 Ceratops montanus
Marsh →USNM 2411；上眼窩角芯、Ceratopsidaeの設立
1889 Ceratops horridus
Marsh →YPM 1820；頭蓋
1889 Triceratops alticornis Marsh →Bison
alticornisに代わって
1889 Triceratops
horridus Marsh →Ceratops horridusに代わって
1889 Triceratops
flabellatus Marsh →YPM 1821；頭蓋
1889 Triceratops galeus
Marsh →情報求む
1889 Monoclonius fissus Cope →AMNH 3998のM.
crassusに代わって
1889 Monoclonius recurvicornis Cope →情報求む
1889
Monoclonius sphenocerus Cope →AMNH 3989；頭蓋
1890 Ceratopsia
Marsh
1890 Ceratopsidae Marsh
1890 Triceratops sulcatus Marsh →USNM
4276；頭蓋
1890 Triceratops prorsus Marsh →YPM 1822；頭蓋
1890
Triceratops serratus Marsh →YPM 1823；頭蓋
1891 Sterrholophus
flabellatus Marsh →情報求む
1891 Torosaurus latus
Marsh →YPM 1830；頭蓋
1891 Torosaurus gladius Marsh →YPM
1831；頭蓋
1891 Triceratops elatus Marsh →USNM 1201
1898
Triceratops calicornis Marsh →情報求む
1898 Triceratops obustus
Marsh →情報求む
1902 Ceratopsoidea Hay
1902 Monoclonius
canadensis Lambe →
1902 Monoclonius
canadensis Lambe →NMC 0491；不完全な頭蓋
1902 Monoclonius dawsoni
Lambe →NMC 1254；不完全な頭蓋と胴椎
1904 Centrosaurus
apertus Lambe →AMNH 5351；
1905 Diceratops
Hatcher →USNM 2412のT. hatcheriに代わって
1905 Triceratops
brevicornis Hatcher →
1905 Triceratops hatcheri Lull →USNM
2412
1913 Styracosaurus albertensis
Lambe →AMNH 5372
1914 Anchiceratops ornatus
Brown →AMNH 5251
1914 Brachiceratops
montanus Gilmore →USNM 7951
1914 Leptoceratops gracillis Brown →
1914 Monoclonius
flexus Brown
1914 Protorosaurus
belli Lambe →NMC 2245とNMC 0491をCeratops belliに代わって
1914 Chasmosaurus belli
Lambe →Protorosaurusに先取権が発生してたため
1915 Centrosaurinae Lambe
1915
Chasmosaurinae Lambe
1915 Eoceratops
canadensis Lambe →Chasmosaurus canadensisに代わって
1915
Triceratops ingens Lull
1917 Monoclonius cutleri Brown
1917
Monoclonius nasicornis Brown
1918 Notoceratops bonarelli
Tapia
1920 Styracosaurus ovatus
Gilmore
1923 Pentaceratops sternbergii Osborn
1923 Protoceratops andrewsi Granger & Gregory
1923
Protoiguanodon mongoliensis Osborn
1923 Psittacosauria Osborn
1923
Psittacosauridae Osborn
1923 Psittacosaurus
mongoliensis Osborn
1924 Anchiceratops longirostris
Sternberg
1925 Arrhinoceratops brachyops
Parks
1930 Pentaceratops
fenestratus Wiman
1931 Psittacosaurus osborni Young
1931
Psittacosaurus tingi Young
1933 Chasmosaurus
brevirostris Lull →ROM 839
1933 Chasmosaurus
kaiseni Brown →AMNH 5401
1933 Triceratops maximus
Brown
1935 Triceratops eurycephalus Schlaikjer
1937
Styracosaurus parksi Brown & Schlaikjer
1940 Centrosaurus
longirostris Sternberg
1940 Chasmosaurus
russelli Sternberg →NMC 8800
1940 Monoclonius lowei
Sternberg
1942 Leptoceratops cerorhynchus Brown &
Sclaikjer
1942 Montanoceratops cerorhynchus
Brown & Sclaikjer
1946 Arrhinoceratops utahensis Gilmore
1946
Torosaurus utahensis Gilmore
1949 Triceratops albertensis
Sternberg
1950 Pachyrhinosaurus canadensis
Sternberg
1953 Microceratops gobiensis Bohlin
1953
Microceratops sulcidens Bohlin
1958 Psittacosaurus
protoiguanodonensis Young
1958 Psittacosaurus
sinensis Young
1962 Psittacosaurus youngi Chao
1975
Bagaceratops rozhdestvensky Maryanska &
Osmolska
1975 Breviceratops kozlowskii
Maryanska & Osmolska
1975 Protoceratops kozlowskii
Maryanska & Osmolska
1983 Psittacosaurus guyangensis
Cheng
1986 Avaceratops lammersi
Dodson
1986 Coronosauria Sereno
1987 Ugrosaurus olsoni Cobabe
& Fastovsky
1988 Psittacosaurus
meileyingensis Sereno, Chao, Cheng & Rao
1988 Psittacosaurus xinjiangensis Sereno & Zhao
1989
Asiaceratops salsopaludalis Nessov &
Kaznyshkina
1989 Chasmosaurus
mariskalensis Lehman
1989 Eucentrosaurus apertus Chure
& McIntosh
1989 Turanoceratops
tardabilis Nessov, Kaznyshkina & Cherepanov
1991 Neoceratopsia
Sereno
1992 Psittacosaurus sattayaraki Buffetaut &
Suteethorn
1992 Udanoceratops tschizhovi Kurzanov
1994 Achelousaurus horneri Sampson
1994 Einiosaurus procurvicornis Sampson
1995
Kulceratops kulensis Nessov
1995 Pachyrhinosaurini
1996
Psittacosaurus mazongshanensis Xu
1996 Psittacosaursu neimongoliensis Russel &
Zhao
1996 Psittacosaurus ordosensis Russel
& Zhao
1997 Archaeoceratops oshimai
Dong & Azuma
1998 Zuniceratops
christopheri Wolfe & Kirtland
1999 Chaoyangsaurus youngi Zhao, Cheng & Xu
2000
Graciliceratops mongoliensis Sereno
2000
Psittacosaurus sibiricus Voronkevich &
Averianov
2001 Chasmosaurus irvinensis Holmes, Forster, Ryan
& Shepherd
2002 Liaoceratops
yanzigouensis Xu, Makovichy, Wang, Norell & You
2003
Bagaceratopidae Alifanov
2003 Bainoceratops efremovi Tereschenko &
Alifanov
2003 Hongshanosaurus houi You, Xu & Wang
2003
Lamaceratops tereschenkoi Alifanov
2003 Magnirostris dodsoni
You & Dong
2003 Platyceratops tatarinovi Alifanov
2003
Serendipaceratops arthurcclarkei Rich & Rich
2005 Centrosaurus brinkmani Ryan and Russell
2005 Auroraceratops rugosus You, Li, Ji, Lamanna, and Dodson
2006 Agujaceratops mariscalensis Lucas, Sullivan and Hunt
2006 Yamaceratops dorngobiensis Makovicky and Norell
2006 Yinlong dowsoni Xu, Forster, Clark and Mo
2006 Psittacosaurus lujiatunensis Zhou, Gao, Fox and Chen
2007 Albertaceratops nesmoi Ryan
Leptoceratopsidae?
Pachyrhinosauridae?
Protoceratopsidae?
Ceratopsomorpha?
Centrosaurini?
Eucentrosaurinae?
Chasmosaurini?
Achelousaurus
Achelousaurus horneri
Agathaumas
sylvestris
Agujaceratops mariscalensis
Agujaceratops
Albertaceratops nesmoi
Anchiceratops
Anchiceratops longirostris
Anchiceratops
ornatus
Arrhinoceratops
Arrhinoceratops brachyops
Arrhinoceratops
utahensis
Asiaceratops
Asiaceratops
salsopaludalis
Auroraceratops
Auroraceratops rugosus
Avaceratops
Avaceratops
lammersi
Bagaceratops
Bagaceratops rozhdestvenskyi
Bainoceratops
efremovi
Bison alticornis
Brachyceratops
Brachyceratops
montaninsis
Breviceratops
Breviceratops
kozlowskii
Centrosaurinae
Centrosaurini
Centrosaurus
Centrosaurus
apertus
Centrosaurus brinkmani
Centrosaurus longirostris
Ceratops horridus
Ceratops montanus
Ceratopsia
Ceratopsidae
Ceratopsoidea
Ceratopsomorha
Chaoyangsaurus
youngi
Chasmosaurinae
Chasmosaurini
Chasmosaurus
Chasmosaurus
belli
Chasmosaurus
brevirostris
Chasmosaurus
canadensis
Chasmosaurus
irvinensis
Chasmosaurus
kaiseni
Chasmosaurus
mariscalensis
Chasmosaurus
russelli
Coronosauria
Diceratops
Diceratops hatcheri
Dysganus
bicarinatus
Dysganus encaustus
Dysganus haydenianus
Dysganus
peiganus
Einiosaurus
Einiosaurus procurvicornis
Eoceratops
Eoceratops
canadensis
Eucentrosaurinae
Eucentrosaurus
Graciliceratops
Graciliceratops mongoliensis
Hongshanosaurus
houi
Kulceratops
Lamaceratops
tereschenkoi
Leptoceratopsidae
Leptoceratops
Leptoceratops
cerorhynchus
Leptoceratops
gracillis
Leptoceratopssp.
Liaoceratops yanzigouensis
Magnirostris dodsoni
Microceratops
Microceratops
gobiensis
Microceratops sulcidens
Monoclonius
Monoclonius
belli
Monoclonius
canadensis
Monoclonius crassus
Monoclonius cutleri
Monoclonius
dawsoni
Monoclonius fissus
Monoclonius flexus
Monoclonius
lowei
Monoclonius nasicornus
Monoclonius recurvicornis
Monoclonius
sphenocerus
Montanoceratops
Montanoceratops
cerorhynchus
Neoceratopsia
Notoceratops
bonarelli
Pachyrhinosauridae
Pachyrhinosaurini
Pachyrhinosaurus
Pachyrhinosaurus
canadensis
Pentaceratops
Pentaceratops
fenestratus
Pentaceratops
sternbergii
Platyceratops tatarinovi
Polyonax
mortuarius
Protiguanodon
Protiguanodon
mongoliensis
Protoceratopsidae
Protoceratops
Protoceratops
andrewsi
Protoceratops hellenikorhinus
Protoceratops kozlowskii
Protorosaurus
Psittacosauria
Psittacosauridae
Psittacosaurus
Psittacosaurus
guyangensis
Psittacosaurus lujiatunensis
Psittacosaurus mazongshanensis
Psittacosaurus
meileyingensis
Psittacosaurus mongoliensis
Psittacosaurus
neimongoliensis
Psittacosaurus ordosensis
Psittacosaurus
osborni
Psittacosaurus protiguanodonensis
Psittacosaurus
sattayaraki
Psittacosaurus sinensis
Psittacosaurus tingi
Psittacosaurus
xinjiangensis
Psittacosaurus youngi
Serendipaceratops
arthurcclarkei
Sterrholophus
Styracosaurus
Styracosaurus
albertensis
Styracosaurus ovatus
Styracosaurus
parksi
Torosaurus
Torosaurus gladius
Torosaurus latus
Torosaurus
utahensis
Triceratops
Triceratops albertensis
Triceratops
alticornis
Triceratops brevicornus
Triceratops calicornis
Triceratops
elatus
Triceratops eurycephalus
Triceratops flabellatus
Triceratops
galeus
Triceratops hatcheri
Triceratops horridus
Triceratops
ingens
Triceratops maximus
Triceratops obtusus
Triceratops
prorsus
Triceratops serratus
Triceratops sulcatus
Turanoceratops
tardabilis
Udanoceratops
Udanoceratops tschizhovi
Ugrosaurus
olsoni
Yamaceratops
Yamaceratops dorngobiensis
Yinlong
Yinlong dowsoni
Zuniceratops
Zuniceratops christopheri